Mischievous and rebellious nine-year-old Milo (Seth Green, voice-over by Seth Dusky) is just beginning summer vacation, and his father (Tom Everett Scott) is leaving for a business trip. While Milo wants his summer to be a fun one, his mother (Joan Cusack) assigns him chores and tasks like taking out the trash. At dinnertime, Milo is given broccoli. His mother has a "no broccoli, no TV" rule which Milo cleverly evades by feeding the broccoli to his pet cat. When Milo's mother finds the cat throwing up from the broccoli, she grounds him and sends him to bed early. After a heated disagreement with his mother, Milo wishes that he never had a mom. Later that night, his wish comes true when his mother is abducted by Martians who plan to steal her "momness" to rear their own young. Milo's quest to save his mom involves stowing away on a spaceship, navigating an elaborate, multi-level planet and taking on the alien nation and their leader, the Supervisor (Mindy Sterling). With the help of tech-savvy subterranean-dwelling earthling Gribble (Dan Fogler), his bionic underground pet Two-Cat (Dee Bradley Baker), and rebellious Martian Ki (Elisabeth Harnois), Milo finds his way back to his mom.

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